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    Arkady Lelej

    A b s t r a c t : The species of Mutillidae from Central Europe are keyed, briefly diagnosed, their distribution and host relationships summarized. New synonymy is proposed: Ronisia brutia (PETAGNA 1787) = Mutilla crimeae STRAND 1917,... more
    A b s t r a c t : The species of Mutillidae from Central Europe are keyed, briefly diagnosed, their distribution and host relationships summarized. New synonymy is proposed: Ronisia brutia (PETAGNA 1787) = Mutilla crimeae STRAND 1917, syn.n.; Smicromyrme triangularis (RADOSZKOWSKI 1865) = S. pouzdranensis HOFFER 1936, syn.n. The key in German language is given also. 34 species in 15 genera are treated.
    Twenty-two species in thirteen genera of Mutillidae and two species in one genus of Bradyno- baenidae from Iran are recorded. Three new species: Dentilla osteni sp. n., Smicromyrme standfussorum sp. n. and Trogaspidia tobiasi sp. n. are... more
    Twenty-two species in thirteen genera of Mutillidae and two species in one genus of Bradyno- baenidae from Iran are recorded. Three new species: Dentilla osteni sp. n., Smicromyrme standfussorum sp. n. and Trogaspidia tobiasi sp. n. are described and figured. The genus Eotrogaspidia Lelej and six species: Eotrogaspidia dives (Smith), Ephutomma angustata (Skorikov), Physetopoda portschinskii (Ra- doszkowski), Smicromyrme minuta Lelej, Neotrogaspidia hammeri (Suarez), and Macroocula sinaica (Invrea) are firstly recorded for Iran. New synonymy is proposed: Eotrogaspidia dives (Smith, 1855) (comb. n.) = Mutilla fumipennis Bingham, 1897 (syn. n.) = Mutilla adscripta Nurse, 1903 (syn. n.). New combinations are proposed: Smicromyrme kermanensis (Lelej, 1984) (comb. n.) and Trogaspidia villosa (Fabricius, 1775) (comb. n.).
    Ilyasov, R.A., Takahashi, J.I., Proshchalykin, M.Y., Lelej, A.S., Lee, M.L., Kwon, H.W., Nikolenko, A. G. (2021). First evidence of presence of Varroa underwoodi mites on native Apis cerana colonies in Primorsky Territory of Russia based... more
    Ilyasov, R.A., Takahashi, J.I., Proshchalykin, M.Y., Lelej, A.S., Lee, M.L., Kwon, H.W., Nikolenko, A. G. (2021). First evidence of presence of Varroa underwoodi mites on native Apis cerana colonies in Primorsky Territory of Russia based on COX1 gene. Journal of Apicultural Science 65 (1), 177-186. doi: 10.2478/JAS-2021-0014. Abstract. The species of genus Varroa mites parasitize on the honey bees of genus Apis. Unlike the well-studied V. destructor and V. jacobsoni mites, V. underwoodi remain less known. According to English language publications, the proven V. underwoodi distribution area of A. cerana colonies covers Nepal, South Korea, Indonesia, Papua New Guinea, Vietnam and China, but not Russia even though it had been described morphometrically in Russian language publications in Russia’s Primorsky Territory. According to Vavilov’s law (1920) of a homologous series, all the species of V. underwoodi, V. destructor and V. jacobsoni have the ability to spill over onto new hosts. ...
    The genus Haba Semenov, 25 species and 3 subspecies of Chrysididae are newly recorded from Russia, as well as five new species are described: Pseudo- malus agnolii Rosa, sp. n. (Tver Prov.), Chrysis mokrousovi Rosa, sp. n. (Abkhazia and... more
    The genus Haba Semenov, 25 species and 3 subspecies of Chrysididae are newly recorded from Russia, as well as five new species are described: Pseudo- malus agnolii Rosa, sp. n. (Tver Prov.), Chrysis mokrousovi Rosa, sp. n. (Abkhazia and Krasnodar Terr.), C. vinokurovi Rosa, sp. n. (Stavropol Terr.), C. lyda Rosa, sp. n. (Krasnodar and Stavropol Terr.), C. sooni Rosa, sp. n. (Primorsky Terr.). Hitherto unknown female of Elampus turcmenicus (Linsenmaier, 1968) is described. Currently the fauna of Russian Chrysididae numbers 330 species and 11 subspecies in 23 genera, including current data.
    A list of 12 species in nine genera occurring in Hong Kong is given. The hitherto unknown male of Bischoffitilla lamellata (Mickel, 1933) is described and illustrated. Six species are newly recorded from Hong Kong. The current number of... more
    A list of 12 species in nine genera occurring in Hong Kong is given. The hitherto unknown male of Bischoffitilla lamellata (Mickel, 1933) is described and illustrated. Six species are newly recorded from Hong Kong. The current number of known mutillid species of Hong Kong is doubled from six to 12. 
    A review of the taxanomic, distributional, and ecological data for the thirty two species, across thirteen genera, of ants recorded from the Kuril Islands is given. In addition, new records - collected during the International Kuril... more
    A review of the taxanomic, distributional, and ecological data for the thirty two species, across thirteen genera, of ants recorded from the Kuril Islands is given. In addition, new records - collected during the International Kuril Island Project (IKIP) - of Myrmica ruginodis kotokui, M. kamtschatica, Leptothorax acervorum and Formica lemani are included from the following eight islands of the archipelago: Matua, Raikoke, Shiashkotan, Onekotan, Makanrushi, Antsiferova, Shumshu and Atlasova. Two species, Crematogaster matsumurai Forel and Lasius neoniger Emery, previously reported by Kuwayama (1967) and Collingwood (1962), respectively, have never been confirmed, despite four large expeditions to the islands, and are considered to be doubtful records for the archipelago. A discussion of the biogeographical distribution of the listed taxa concludes.
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    Petersenidia gen. nov. (type species: Smicromyrme fukudai TSUNEKl), hitherto unknown female of Myrmosa eos LELEJ and male of Squamulotilla ex- ilipunclala CHEN are described. Smicromyrme wolani (ZAV.) and Ephutomma... more
    Petersenidia  gen.  nov. (type  species:  Smicromyrme fukudai  TSUNEKl),
    hitherto  unknown  female  of Myrmosa  eos  LELEJ  and  male  of Squamulotilla  ex-
    ilipunclala CHEN  are described.  Smicromyrme  wolani (ZAV.)  and  Ephutomma naja
    (ZAV.) are newly  recorded from Japan and  M.  eos LELEJ  and S. exilipunctata CHEN
    are newly  recorded from Korea.  New synonymy in  two species is  given.
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    Citation: Brothers DJ, Lelej AS (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research 60: 1–97. https://doi. Abstract This study aimed to resolve the... more
    Citation: Brothers DJ, Lelej AS (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research 60: 1–97. https://doi. Abstract This study aimed to resolve the differences in the two currently used classifications of Mutillidae, which differ in many respects. Cladistic analyses of 101 genera and subgenera of Mutillidae (represented by females of 253 species and males of 260 species) and four outgroups (pepsine Pompilidae, anthobos-cine Tiphiidae and both fedtschenkiine and sapygine Sapygidae) based on 230 morphological characters treated in various ways, produced most-parsimonious trees which were in broad agreement but differed in many details. Evaluation of these results led to the proposal of a compromise tree which reflected each proposed taxon as monophyletic, while trying to keep disruptions to the current classifications to a minimum. The result differs from both previous classifications, and proposes the recognition of eight subfamilies: Myrmosinae (with the tribes Kudakrumiini and Myrmosini), Pseudophotopsidinae, Rho-palomutillinae, Ticoplinae (with the tribes Smicromyrmillini and Ticoplini), Sphaeropthalminae (with the tribes Sphaeropthalmini, Dasymutillini trib. n., and Pseudomethocini with the subtribes Euspinoliina subtrib. n. and Pseudomethocina), Myrmillinae, Dasylabrinae (with the tribes Apteromutillini trib. n. and Dasylabrini) and Mutillinae (with the tribes Ctenotillini trib. n., Smicromyrmini, Mutillini with the subtribes Ephutina and Mutillina, and Trogaspidiini). Notably, Myrmosinae were consistently strongly supported as monophyletic with the remaining Mutillidae (disagreeing with a recent molecular analysis), and thus retained as a mutillid subfamily. The placements of all currently valid genera and subgenera in the proposed classification are provided.
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    A review of type specimens of nine nominal taxa, described by E. Eversmann is given. The lectotypes are designated for Scolia grisea Eversmann, 1849, S. sareptana Eversmann, 1849, and S. schrenckii Eversmann, 1846.... more
    A review of type specimens of nine nominal taxa, described by E.
    Eversmann  is  given.  The  lectotypes  are  designated  for  Scolia  grisea  Eversmann,
    1849,  S.  sareptana  Eversmann,  1849,  and  S.  schrenckii  Eversmann,  1846.  The
    neotype is designated for Vespa galbula Pallas, 1771. New synonymy is proposed
    for Colpa galbula (Pallas, 1771), comb. n. = Scolia sexmaculata Fabricius, 1781,
    syn. n., = S. interrupta Fabricius, 1781, syn. n., = S. sareptana Eversmann, 1849,
    syn. n.; Scolia quadricincta Scopoli, 1786, stat. resurr. = S. flaviceps Eversmann,
    1846, syn. n. According to Articles 23.9.1.2, 23.9.2 of Code (ICZN, 1999) Scolia
    fallax  Eversmann,  1849  is  nomen  protectum  and  has  precedence  over  the  older
    name Vespa tricolor Pallas, 1771, nomen oblitum. Checklist of Russian Scoliidae (5
    genera, 20 species) is given.
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    The genus Nipponodipogon Ishikawa, 1965 is newly recorded from China (Guangdong, Hainan, and Yunnan) and the Oriental Region. Two new species, N. orientalis Loktionov, Lelej & Xu, sp. n. (Guangdong, Hainan, Yunnan) and N. shimizui... more
    The genus Nipponodipogon Ishikawa, 1965 is newly recorded from China (Guangdong, Hainan, and Yunnan) and the Oriental Region. Two new species, N. orientalis Loktionov, Lelej & Xu, sp. n. (Guangdong, Hainan, Yunnan) and N. shimizui Loktionov, Lelej & Xu, sp. n. (Guangdong, Yunnan), are described and illustrated. The updated key to the species based on Shimizu et al. (2015) is given.
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    ... Smicromyrme drupa: Mickel, 1933b: 417, male (Taiwan); 1935: 273, male (key); Chen, 1957: 180, 202, male (Taiwan); Tsuneki, 1972: 18, male (Taiwan); 1993: 40, male (Taiwan). MATERIAL (2 males). ... Doi Pui near Chiang Mai, 9. VIII 1988... more
    ... Smicromyrme drupa: Mickel, 1933b: 417, male (Taiwan); 1935: 273, male (key); Chen, 1957: 180, 202, male (Taiwan); Tsuneki, 1972: 18, male (Taiwan); 1993: 40, male (Taiwan). MATERIAL (2 males). ... Doi Pui near Chiang Mai, 9. VIII 1988 (Y. Notsu) [NSMT]. DESCRIPTION. ...
    Eleven species of Promecidia Lelej, 1996 are reviewed and keyed, and the diagnosis of the genus is given. The genus Promecidia is newly recorded from China and Promecidia abnormis Lelej, sp. n. (China: Guangdong, Hainan) and Promecidia... more
    Eleven species of Promecidia Lelej, 1996 are reviewed and keyed, and the diagnosis of the genus is given. The genus Promecidia is newly recorded from China and Promecidia abnormis Lelej, sp. n. (China: Guangdong, Hainan) and Promecidia chui Lelej & Xu, sp. n. (China: Yunnan, Hainan) are described and illustrated. New combination is proposed for Promecidia boopis (Kohl, 1882), comb. n. (from the genus Petersenidia Lelej, 1996). New status is proposed for Promecidia saturnia (Mickel, 1935), stat. n. and Promecidia samawangensis (Mickel, 1935), stat. n.
    Kozlov (1994) established a new parasitic proctotrupoid family, Renyxidae, for a single species, Renyxa incredibilis, from Primorskij Krai, in the Russian Far East. A new genus and species, Hsiufuropronia chaoi, was described from China,... more
    Kozlov (1994) established a new parasitic proctotrupoid family, Renyxidae, for a single species, Renyxa incredibilis, from Primorskij Krai, in the Russian Far East. A new genus and species, Hsiufuropronia chaoi, was described from China, Beijing (Yang 1997) in the family Roproniidae. Lelej and Kozlov (1999) noticed that Renyxa was preoccupied and was initially used by Kurochkin and Slankis (1973) for a genus of flatworms (Cestoda: Litobothridae). According to Articles 39, 60 of the Code (ICZN 1999), the names of the genus and family were changed to Proctorenyxa Lelej & Kozlov, 1999 (Lelej & Kozlov 1999). He et al. (2002) transferred Hsiufuropronia from Roproniidae to Proctorenyxidae based on Yang's original description and illustrations, and synonymized Hsiufuropronia Yang under Proctorenyxa Lelej & Kozlov. Later, He and Xu (2015) proposed that the family name Proctorenyxidae should be replaced by Hsiufuroproniidae and the genera Proctonenyxa Lelej & Kozlov and Hsiufuropronia Yang should be separate. However, the replacement name Hsiufuroproniidae contradicts article 39 of the Code (ICZN 1999).
    ... ETYMOLOGY. The species is dedicated to Vladimir Veniaminovich Popov, who was an expert in Apoidea and revised the genus Biastes. Key to the species of Biastes of the Russian Far East (males) ... Storozhenko, principal investigator).... more
    ... ETYMOLOGY. The species is dedicated to Vladimir Veniaminovich Popov, who was an expert in Apoidea and revised the genus Biastes. Key to the species of Biastes of the Russian Far East (males) ... Storozhenko, principal investigator). FAMILY TIPULIDAE ...
    Phylogenetic analysis of fifteen higher taxa (subfamilies and tribes) with hypothetical ancestor as outgroup reveal the relationships: Ancestor + (Myrmosinae + (Kudakrumiinae + {Pseudophotopsidinae + (Ticoplinae + ({Rhopalomutillinae +... more
    Phylogenetic analysis of fifteen higher taxa (subfamilies and tribes) with hypothetical ancestor as outgroup reveal the relationships: Ancestor + (Myrmosinae + (Kudakrumiinae + {Pseudophotopsidinae + (Ticoplinae + ({Rhopalomutillinae + ((Ephutinae + Dasylabrinae) + Sphaeropthalminae)} + (Myrmillinae + Mutillinae)))})). A tree generated from fifteen higher taxa and hypothetical ancestor based on 71 "best" from 89 analysed characters by PAUP yielded a
    ARKADY S. LELEJ1 & EROL YILDIRIM2 1Institute of Biology and Soil Science, Far Eastern Branch of Russian Academy of Sciences, Vladivostok-22, 690022, Russia. E-mail: lelej@biosoil.ru 2Atatürk University, Faculty of... more
    ARKADY S. LELEJ1 & EROL YILDIRIM2 1Institute of Biology and Soil Science, Far Eastern Branch of Russian Academy of Sciences, Vladivostok-22, 690022, Russia. E-mail: lelej@biosoil.ru 2Atatürk University, Faculty of Agriculture, Department of Plant Protection, 25240, Erzurum, Turkey. ...
    Keys to 55 genera of spider wasps of Russia and neighbouring countries in females and males are given. Of them 34 genera are distributed in Russia. An annotated list of genera with type species and distribution data within Russia and... more
    Keys to 55 genera of spider wasps of Russia and neighbouring countries in females and males are given. Of them 34 genera are distributed in Russia. An annotated list of genera with type species and distribution data within Russia and biogeographical regions is given. The genus Xenaporus Ashmead, 1902 and X. eremocanus Wolf, 1990 are newly recorded from Russia. According to ICZN 1995 (Opinion 1820) new synonymy (valid name first) is proposed for the type species of genus Cryptocheilus Panzer, 1806: Sphex annulata Fabricius, 1798 (=Pompilus alternatus Lepeletier de Saint Fargeau, 1845, syn. nov.; =Pompilus comparatus Smith, 1855, syn. nov.; =Priocnemis culpabilis Costa, 1893, syn. nov.; Salius annulatilis Richards, 1935, syn. nov.).

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