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| status = G5
| status_system = TNC
| status_ref = <ref>{{cite iucn |author=Beckman, E. |date=2016 |title=''Quercus agrifolia'' |volume=2016 |page=e.T194049A2295175 |doi=10.2305/IUCN.UK.2016-1.RLTS.T194049A2295175.en |access-date=12 November 2021}}</ref>
| genus = Quercus
| display_parents = 2
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| synonyms_ref = <ref>{{ThePlantList |id=kew-171483 |taxon=Quercus agrifolia |authority=Née}}</ref>
| synonyms = {{collapsible list|bullets = true
| ''Quercus acroglandis'' <small>Kellogg</small>
| ''Quercus acutiglandis'' <small>Sarg.</small>
| ''Quercus agrifolia'' var. ''frutescens'' <small>Engelm.</small>
| ''Quercus agrifolia'' var. ''oxyadenia'' <small>(Torr.) J.T.Howell</small>
| ''Quercus oxyadenia'' <small>Torr.</small>
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}}
'''''Quercus agrifolia''''', the '''California live oak''',<ref>{{PLANTS|id=QUAG|taxon=Quercus agrifolia|access-date=30 July 2015}}</ref> or '''coast live
This species is commonly [[sympatric]] with [[canyon live oak]] (''Q.
==Description==
Coast live oak typically has a much-branched trunk and reaches a mature height of {{convert|10|–|25|m|ft|sp=us}}. Some specimens may attain an age exceeding 1000 years. Examples of this include the Grand Oak of Cherry Valley, California,<ref>{{Cite web|url=https://www.monumentaltrees.com/en/usa/california/riversidecounty/8948_highlandspringsresort/17622/|title=Coast live oak 'The Grand Oak' at Highland Springs Resort, Cherry Valley, California, United States}}</ref> the [[Encino Oak Tree]], which died in the 1990s (part of the stump has been preserved)<ref>{{Cite web|url=http://www.highbeam.com/doc/1G1-83858687.html|archive-url = https://web.archive.org/web/20121021220556/http://www.highbeam.com/doc/1G1-83858687.html|archive-date = 2012-10-21|title = Questia}}</ref> and the Pechanga Great Oak.<ref name="pechanga-nsn.gov">{{Cite web|url=https://www.pechanga-nsn.gov/index.php/history/the-great-oak|title=Pechanga Band of Luiseño Indians - the Great Oak}}</ref>▼
▲Coast live oak typically has a much-branched trunk and reaches a mature height of {{convert|10|–|25|m|ft|
The trunk, particularly for older individuals, may be highly contorted, massive and gnarled. The crown is broadly rounded and dense, especially when aged 20 to 70 years; in later life the trunk and branches are more well defined and the leaf density lower.<ref name=FNA/> The oldest specimens might exceed 20 feet in trunk circumference and 100 feet height.<ref name="pechanga-nsn.gov"/><ref>{{Cite web|url=https://www.lindaymakutadds.com/the-lang-oak-encino-oak-tree/|title = The Great Encino Oak Tree}}</ref>▼
▲The trunk, particularly for older individuals, may be highly contorted, massive and gnarled. The crown is broadly rounded and dense, especially when aged 20 to 70 years; in later life the trunk and branches are more well defined and the leaf density lower.<ref name=FNA/> The oldest specimens might exceed {{Convert|20
The [[leaf|leaves]] are dark green, oval, often [[wikt:convex|convex]] in shape, {{convert|2|–|7|cm|in|abbr=on}} long and {{convert|1|–|4|cm|in|abbr=on}} broad; the leaf margin is spiny-toothed (spinose), with sharp thistly fibers that extend from the lateral leaf veins. The outer layers of leaves are designed for maximum [[sun|solar]] absorption, containing two to three layers of [[photosynthetic]] cells.<ref name=FNA/>▼
▲The [[
These outer leaves are deemed to be small in size to more efficiently re-radiate the heat gained from solar capture. Shaded leaves are generally broader and thinner, having only a single layer of photosynthetic cells. The convex leaf shape may be useful for interior leaves which depend on capturing reflected light scattered in random directions from the outer canopy.<ref name=FNA>{{eFloras|1|233501005|Quercus agrifolia |first=Kevin C. |last=Nixon |volume=3}}</ref>▼
▲These outer leaves are deemed to be small in size to more efficiently re-radiate the heat gained from solar capture. Shaded leaves are generally broader and thinner, having only a single layer of photosynthetic cells. The convex leaf shape may be useful for interior leaves which depend on capturing reflected light scattered in random directions from the outer canopy.<ref name=FNA>{{eFloras|1|233501005|Quercus agrifolia |first=Kevin C. |last=Nixon |volume=3}}</ref> The leaf shape may be also useful as condensation surfaces for "dew and mist, which would allow the tree to survive years with limited rainfall".<ref>{{Cite web |title=Coastal live oak - Quercus agrifolia |url=https://smmtc.org/plantofthemonth/Coastal_Live_Oak.php |access-date=2023-11-10 |website=Santa Monica Mountains Trails Council |language=en}}</ref>
[[File:cloacorne.jpg|thumb|left|Acorns and leaves]]▼
The [[flower]]s are produced in early-to-mid spring; the male flowers are pendulous [[catkin]]s {{convert|5|–|10|cm|in|abbr=on|frac=4}} long, the female flowers inconspicuous, less than {{convert|0.5|cm|in|abbr=on|frac=4}} long, with 1–3 clustered together. The [[fruit]] is a slender reddish brown [[acorn]] {{convert|2|-|3.5|cm|in|abbr=on|frac=4}} long and {{convert|1|-|1.5|cm|in|abbr=on|frac=8}} broad, with the basal quarter enclosed in a [[Calybium and cupule|cupule]]; unusually for a red oak, the acorns mature about 7–8 months after pollination (most red oak acorns take 18 months to mature).<ref name=FNA/>
===Recognized varieties===
There are two [[
{| class="wikitable"
|-
! Image !! Scientific name!! Description !! Distribution
|-
|[[File:Quercus agrifolia leaves and acorn.jpg|120px]] || ''Quercus agrifolia'' var. ''agrifolia'' ||Leaves that are glabrous to slightly hairy on the abaxial side, especially near the leaf vein axils. Hybrids with ''[[Quercus kelloggii|Q.
|-
| ||''Quercus agrifolia'' var. ''oxyadenia'' ||
|-
|}
===Hybridity===
Several hybrids between coast live oak and other red oak species have been documented. Hybrids with [[interior live oak]] (''Q.
==Etymology==
In naming the species, [[Luis Née|Née]] compared it to a species illustrated in [[Leonard Plukenet]]
==
[[File:CoastLiveoakSLOCountyCA.jpg|alt=Tree growing by a roadside in California|thumb|Coast live oak, [[San Luis Obispo County, California
Coast live oak is the only California native oak that actually thrives in the coastal environment, although it is rare on the immediate shore; it enjoys the mild winter and summer climate afforded by ocean proximity, and it is somewhat tolerant of aerosol-borne sea salt. The coastal fog supplies relief from the rainless California summer heat.▼
Normally the tree is found on well
▲
It is the dominant overstory plant of the coast live oak [[woodland]] habitat, often joined by [[California bay laurel]] and [[California buckeye]] north of [[Big Sur]]. Associated understory plants include [[toyon]], various [[manzanita]]s and [[western poison-oak]].
== Ecology ==
▲Normally the tree is found on well drained soils of coastal hills and plains, often near year round or [[perennial stream]]s. It may be found in several natural communities including coast live oak woodland, [[Engelmann oak]] woodland, [[valley oak]] woodland and both northern and southern mixed evergreen forests. While normally found within {{convert|100|km|mi|abbr=off|sp=us}} of the [[Pacific Ocean]] at elevations less than {{convert|700|m|ft|abbr=off|sp=us}}, in southern California it occasionally occurs at up to {{convert|1,500|m|ft|abbr=off|sp=us}} in altitude.
{{multiple image
| border =
| total_width = 350
| image_style = border:none;
| perrow = 1/2/2/2/2
| image1 = Quercus agrifolia coast live oak and Toxicodendron diversilobum pacific poison oak.jpg
|image2 = Viscid bolete imported from iNaturalist photo 248008629 on 8 November 2023.jpg
| image3 = Oak Titmouse (40414129682).jpg
|image4 = California Sister imported from iNaturalist photo 17311317 on 8 November 2023.jpg
| image5= Live Oak Apple Gall Wasp imported from iNaturalist photo 117933461 on 8 November 2023.jpg
|image6= Acorn Woodpecker imported from iNaturalist photo 114896966 on 8 November 2023.jpg
|image7 = Lace lichen imported from iNaturalist photo 326308289 on 8 November 2023.jpg
| image8 = Live Oak Witches' Broom Fungus imported from iNaturalist photo 354032235 on 27 February 2024.jpg
| image9 = Bewick's Wren (53396677802).jpg
| footer = [[Pacific poison oak]], [[Aureoboletus flaviporus|viscid bolete]], [[oak titmouse]], [[California sister]], [[Amphibolips quercuspomiformis|live-oak apple]], [[acorn woodpecker]], [[lace lichen]], [[live oak witch's broom]], [[Bewick's wren]]
}}
The California oak moth (''[[Phryganidia californica]]'') caterpillar subsists entirely on living and fallen leaves of the Coast Live Oak. In 8–10 year cycles, the caterpillar will appear in sufficient abundance to denude healthy trees. The trees recover, and botanists speculate that the species provide mutual benefit, possibly in the form of fertilizer for the oak.<ref>{{cite web
Live oak trees, among other western oaks, are also known to support acorn woodpeckers, which store their acorns in tree trunks and remove them when they want to eat. <ref> {{cite web |title=An Enduring Oak Mystery: Synchronized Acorn Booms |url=https://baynature.org/article/an-enduring-oak-mystery-synchronized-acorn-booms/}}</ref>
▲The California oak moth (''[[Phryganidia californica]]'') caterpillar subsists entirely on living and fallen leaves of the Coast Live Oak. In 8–10 year cycles, the caterpillar will appear in sufficient abundance to denude healthy trees. The trees recover, and botanists speculate that the species provide mutual benefit, possibly in the form of fertilizer for the oak.<ref>{{cite web|url=http://trees.stanford.edu/ENCYC/QUERCUS.htm|title=Quercus (Oak) Notes – Trees of Stanford & Environs|website=trees.stanford.edu}}</ref> The coast live oak is also the only known foodplant of ''[[Chionodes vanduzeei]]'' [[caterpillar]]s.
== Allergenicity ==
The pollen of the coast live oak is a severe allergen. Pollination occurs in spring.<ref name="pollen">{{Cite web|url=http://www.pollenlibrary.com/Local/Specie/Quercus+agrifolia/in/Santa%20clara%20County/CA/|title = Coastal Live Oak (Quercus agrifolia) Species Details and Allergy Info, Santa clara county, California}}</ref>
==
===Historical
{{original research section|date=May 2024}}
[[File:Los Encinos live oak.jpg|thumb|upright=1.
At least twelve distinct cultures of [[Native Americans in the United States|Native Americans]] are known to have consumed the acorns as a dietary staple.<ref>{{cite book |last1=Moerman |first1=Daniel |title=Native American Food Plants: An Ethnobotanical Dictionary |date=2010 |publisher=Timber Press |pages=472–473}}</ref> The seeds were ground into meal, which after being washed was boiled into mush or baked in ashes to make bread.<ref>{{cite book |last=Whitney |first=Stephen |title=Western Forests (The Audubon Society Nature Guides) |date=1985 |publisher=Knopf |location=New York |isbn=0-394-73127-1 |page=[https://archive.org/details/westernforests00whit/page/383 383] |url=https://archive.org/details/westernforests00whit/page/383 }}</ref> In the 18th century, [[Spanish colonization of the Americas|Spaniards]] in the [[San Fernando Valley]] used the wood for charcoal to fire [[kiln]]s in making [[adobe]]. Later this form of charcoal would be utilized in the baking, [[gunpowder]] and [[electric power]] industries.▼
▲At least twelve distinct cultures of [[Native Americans in the United States|Native Americans]] are known to have consumed the acorns as a dietary staple.
In the 18th and 19th centuries [[shipbuilder]]s sought out the odd angular branches to make special joints. [[Settler|Pioneer]]s moving west would harvest small amounts for making farm implements and wagon wheels, but the greatest impact was the wholesale clearing of oak woodlands to erect sprawling cities such as [[San Diego]] and [[San Francisco]]. The irregular shape often let the tree escape widespread harvest for building timbers, and also led the early settlers to endow the coast live oak with mystical qualities. Its stateliness has made it a subject of historical [[Landscape art|landscape painter]]s throughout California modern history since the mid-19th century.
===Modern
Coast live oak has also become a common addition to western
Also, if incorporating it into a landscaping scheme with artificial irrigation, it is important to avoid regular watering within the oak's [[Tree canopy|drip line]] (canopy), since wet soil in the summer increases infection rates by soil-borne ''[[Phytophthora]]'' diseases like [[sudden oak death]].<ref name="Sudden Oak Death: Management Strategies">{{cite web|url=http://www.plantmanagementnetwork.org/php/shared/sod/|title=Sudden Oak Death and Associated Diseases Caused by Phytophthora ramorum|last=J. M. Davidson|date=7 July 2003|publisher=Plant Management Network|access-date=12 January 2010}}</ref>
==Culture==
The coast live oak, especially in its [[Spanish language|Spanish]] forms ''encino'' or ''encina'', ''encinitas'' "little oaks", and ''encinal'' "oak grove", gave its name to seven land grants across California and to many communities and geographic features. These include [[Rancho Los Encinos]], the Los Angeles community of [[Encino, Los Angeles|Encino]], [[Encinitas, California|Encinitas]] near [[San Diego, California|San Diego]], and Encinal del Temescal, now the city of [[Oakland, California|Oakland]].<ref>Gudde, Erwin, and William Bright, ''California Place Names'', University of California Press, 4th edition, 1998, {{ISBN|
[[Paso Robles]] (fully 'El Paso de Robles' or 'Pass of the Oaks') also refers to the ubiquitous live oaks in the region as a geographical place name.
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==Bibliography==
* {{Cite book|last=Balls|first=Edward Kent|url=https://www.worldcat.org
* C. Michael Hogan, California Arts and Sciences Institute calasinstitute.org (2023) ''Coast Live Oak''. ed. Arthur Dawson. Encyclopedia of Earth. National Council for Science and Environment. Washington DC
* {{Cite book|last1=
* {{Cite book|last1=Sawyer|first1=John O.|title=A Manual of California Vegetation|last2=Keeler-Wolf|first2=Todd|publisher=[[California Native Plant Society]]|year=1995|isbn=978-0943460260|location=|pages=241}}
== External links ==
{{Commons category|Quercus agrifolia}}
* {{NPIN |id=QUAG}}
* [http://www.laspilitas.com/groups/oaks/california_oak1.html Las Pilitas: California Oaks]
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[[Category:Quercus|agrifolia]]
[[Category:Flora of California]]
[[Category:Trees of
[[Category:Natural history of the California chaparral and woodlands]]
[[Category:Natural history of the California Coast Ranges]]
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[[Category:Drought-tolerant trees]]
[[Category:Ornamental trees]]
[[Category:Oaks of Mexico]]
|