Abstract
The innovations of agricultural production and their extensive dispersal promoted the transformation of human livelihoods and profoundly influenced the evolution of human-land relationships in late prehistoric Eurasia. The Steppe and Silk Roads (SSRs) played important roles in the transcontinental exchange and dispersal of cereal crops and livestock related to agricultural innovation across Eurasia before the Han Dynasty (202 BC to AD 220), while the geographical-temporal variations in prehistoric subsistence in relation to the spread and exchange of cereal crops and livestock originating from different areas of Eurasia still remain unclear. In this paper, we explore these issues based on the review and analysis of published archaeobotanical, zooarchaeological, and carbon-stable isotope data from human bones from Neolithic-Early Iron Age sites in areas along the SSRs, with a comparison to updated results based on radiocarbon dating and ancient DNA analyses. Our results suggest that humans engaged in hunting game, while foxtail/broomcorn millet cultivation gradually became the primary subsistence strategy in Eastern SSRs from 10,500 to 6000 a BP. In contemporaneous Western SSRs, humans mainly cultivated wheat/barley and raised sheep/goats, cattle, and pigs. Trans-Eurasian exchange, which is reflected by the mixed utilization of wheat/barley and millet, emerged in the south-central Steppe during 6000–4000 a BP, while millet cultivation and pig husbandry became the dominant livelihoods in most areas of Eastern SSRs. During 4000–2200 a BP, Silk Roads became the major passageway for trans-Eurasian exchange, the interactive development of oasis agriculture and pastoralism facilitated intensive human settlement in the Central Silk Roads, and subsistence strategies substantially changed with significant geographical differences in Eastern SSRs, while subsistence in some areas of Western SSRs was evidently affected by the introduction and adoption of millet crops after 3000 a BP. The geographical-temporal variations in subsistence in the SSRs from the Neolithic to Early Iron Age were primarily affected by the prehistoric dispersal of farming groups across Eurasia, which was accompanied by the spread of cereal crops/livestock, while the impacts of climate change still need to be further evaluated.
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Allentoft M E, Sikora M, Sjögren K G, Rasmussen S, Rasmussen M, Stenderup J, Damgaard P B, Schroeder H, Ahlström T, Vinner L, Malaspinas A S, Margaryan A, Higham T, Chivall D, Lynnerup N, Harvig L, Baron J, Della Casa P, Dabrowski P, Duffy P R, Ebel A V, Epimakhov A, Frei K, Furmanek M, Gralak T, Gromov A, Gronkiewicz S, Grupe G, Hajdu T, Jarysz R, Khartanovich V, Khokhlov A, Kiss V, Kolář J, Kriiska A, Lasak I, Longhi C, McGlynn G, Merkevicius A, Merkyte I, Metspalu M, Mkrtchyan R, Moiseyev V, Paja L, Pálfi G, Pokutta D, Pospieszny L, Price T D, Saag L, Sablin M, Shishlina N, Smrcka V, Soenov V I, Szeverényi V, Tóth G, Trifanova S V, Varul L, Vicze M, Yepiskoposyan L, Zhitenev V, Orlando L, Sicheritz-Pontén T, Brunak S, Nielsen R, Kristiansen K, Willerslev E. 2015. Population genomics of Bronze Age Eurasia. Nature, 522: 167–172
Alley R, Ágústsdóttir A. 2005. The 8k event: Cause and consequences of a major Holocene abrupt climate change. Quat Sci Rev, 24: 1123–1149
An C B, Wang W, Duan F T, Huang W, Chen F H. 2017. Environmental changes and cultural exchange between East and West along the Silk Road in arid Central Asia (in Chinese). Acta Geogr Sin, 72: 875–891
Arbuckle B S, Kansa S W, Kansa E, Orton D, Çakırlar C, Gourichon L, Atici L, Galik A, Marciniak A, Mulville J, Buitenhuis H, Carruthers D, de Cupere B, Demirergi A, Frame S, Helmer D, Martin L, Peters J, Pöllath N, Pawlowska K, Russell N, Twiss K, Würtenberger D. 2014. Data sharing reveals complexity in the westward spread of domestic animals across Neolithic Turkey. PLoS ONE, 9: e99845
Atici L, Pilaar Birch S E, Erdoğu B. 2017. Spread of domestic animals across Neolithic western Anatolia: New zooarchaeological evidence from Ugurlu Höyük, the island of Gökçeada, Turkey. PLoS ONE, 12: e0186519
Balasse M, Gillis R, Živaljević I, Berthon R, Kovačiková L, Fiorillo D, Arbogast R M, Bălăşescu A, Bréhard S, Nyerges É Á, Dimitrijević V, Bánffy E, Domboróczki L, Marciniak A, Oross K, Vostrovská I, Roffet-Salque M, Stefanović S, Ivanova M. 2021. Seasonal calving in European Prehistoric cattle and its impacts on milk availability and cheese-making. Sci Rep, 11: 8185
Bellwood P. 2005. First Farmers: The Origins of Agricultural Societies. Cornwall: Black Publishing
Bendrey R. 2011. Some like it hot: Environmental determinism and the pastoral economies of the later prehistoric Eurasian steppe. Pastor Res Policy Pract, 1: 8
Benecke N, von den Driesch A. 2003. Horse exploitation in the Kazakh Steppes during the Eneolithic and Bronze Age. In: Levine M, Renfrew C, Boyle K, eds. Prehistoric Steppe Adaptation and the Horse. Cambridge: McDonald Institute for Archaeological Research. 69–82
Bocherens H, Mashkour M, Billiou D. 2000. Palaeoenvironmental and archaeological implications of isotopic analyses (13C, 15N) from Neolithic to Present in Qazvin Plain (Iran). Environ Archaeol, 5: 1–19
Boivin N, Fuller D Q, Crowther A. 2012. Old World globalization and the Columbian exchange: Comparison and contrast. World Archaeol, 44: 452–469
Bradley D G, Loftus R T, Cunningham P, MacHugh D E. 1998. Genetics and domestic cattle origins. Evol Anthropol, 6: 79–86
Brunson K, He N, Dai X. 2016. Sheep, cattle, and specialization: New zooarchaeological perspective on the Taosi Longshan. Int J Osteoarchaeol, 26: 460–475
Budd C, Lillie M, Alpaslan-Roodenberg S, Karul N, Pinhasi R. 2013. Stable isotope analysis of Neolithic and Chalcolithic populations from Aktopraklik, northern Anatolia. J Archaeol Sci, 40: 860–867
Chechushkov I V, Kosintsev P A. 2020. The Botai horse practices represent the neolithization process in the central Eurasian steppes: Important findings from a new study on ancient horse DNA. J Archaeol Sci-Rep, 32: 102426
Chen F H, Dong G H, Zhang D J, Liu X Y, Jia X, An C B, Ma M M, Xie Y W, Barton L, Ren X Y, Zhao Z J, Wu X H, Jones M K. 2015. Agriculture facilitated permanent human occupation of the Tibetan Plateau after 3600 B.P. Science, 347: 248–250
Chen F H, Yu Z C, Yang M L, Ito E, Wang S M, Madsen D B, Huang X Z, Zhao Y, Sato T, Birks H J B, Boomer I, Chen J H, An C B, Wünnemann B. 2008. Holocene moisture evolution in arid central Asia and its out-of-phase relationship with Asian monsoon history. Quat Sci Rev, 27: 351–364
Chen T T, Qiu M H, Liu R L, Li H M, Hou H W, Howarth P, Bowring S, Zhou A F. 2020. Human responses to climate change in the late prehistoric Western Loess Plateau, Northwest China. Radiocarbon, 62: 1193–1207
Chen X L, Fang Y M, Hu Y W, Hou Y F, Lv P, Song G D, Yuan J, Richards M P. 2017. Stable carbon and nitrogen analysis of humans and animals to imply the complexity of prehistoric economic lives: A case study at Wadian site of Yu City (in Chinese). Huaxia Archaeol, 4: 70–79
Chen X X, Yu S Y, Wang Q Z, Cui X X, Underhill A P. 2020. More direct evidence for early dispersal of bread wheat to the eastern Chinese coast ca. 2460–2210 BC. Archaeol Anthropol Sci, 12: 233
Childe V G. 1930. The Bronze Age. Cambridge: Cambridge University Press
Colledge S, Conolly J, Shennan S. 2004. Archaeobotanical evidence for the spread of farming in the Eastern Mediterranean. Curr Anthropol, 45: S35–S58
Cramp L J E, Ethier J, Urem-Kotsou D, Bonsall C, Borić D, Boroneanţ A, Evershed R P, Perić S, Roffet-Salque M, Whelton H L, Ivanova M. 2019. Regional diversity in subsistence among early farmers in Southeast Europe revealed by archaeological organic residues. Proc R Soc B, 286: 20182347
Crawford G, Underhill A, Zhao Z J, Lee G A, Feinman G, Nicholas L, Luan F S, Yu H G, Fang H, Cai F S. 2005. Late Neolithic plant remains from Northern China: Preliminary results from Liangchengzhen, Shandong. Curr Anthropol, 46: 309–317
Cui Y P, Hu Y W, Chen H H, Dong Y, Guan L, Weng Y, Wang C S. 2006. Stable isotopic analysis on human bones from Zongri site (in Chinese). Quat Sci, 26: 604–611
Curry A. 2013. Archaeology: The milk revolution. Nature, 500: 20–22
Dai L L, Li Z P, Hu Y W, Zhao C Q, Wang C S. 2014. The kill-off pattern of the Caprids and the animal resource utilization strategy of Xinzhai site (in Chinese). Archaeology, 1: 94–103
d’Alpoim Guedes J, Bocinsky R K. 2018. Climate change stimulated agricultural innovation and exchange across Asia. Sci Adv, 4: eaar4491
Dani A H, Masson V M. 2002. History of Civilizations of Central Asia·I (Chinese Edition) (in Chinese). Beijing: China Translation and Publishing Corporation. 86–88, 118–176, 352–363
de Cupere B, Duru R. 2003. Faunal remains from Neolithic Höyücek (SW-Turkey) and the presence of early domestic cattle in Anatolia. Paléorient, 29: 107–120
Decaix A, Messager E, Tengberg M, Neef R, Lyonnet B, Guliyev F. 2016. Vegetation and plant exploitation at Mentesh Tepe (Azerbaijan), 6th-3rd millennium BC initial results of the archaeobotanical study. Quat Int, 395: 19–30
Deng Z H, Fuller D Q, Chu X L, Cao Y P, Jiang Y C, Wang L Z, Lu H Y. 2019. Assessing the occurrence and status of wheat in late Neolithic central China: The importance of direct AMS radiocarbon dates from Xiazhai. Veget Hist Archaeobot, 29: 61–73
Diamond J, Bellwood P. 2003. Farmers and their languages: The first expansions. Science, 300: 597–603
Dodson J R, Li X Q, Zhou X Y, Zhao K L, Sun N, Atahan P. 2013. Origin and spread of wheat in China. Quat Sci Rev, 72: 108–111
Dong G H, Du L Y, Wei W Y. 2020b. The impact of early trans-Eurasian exchange on animal utilization in northern China during 5000–2500 BP. Holocene, 31: 294–301
Dong G H, Li R, Lu M X, Zhang D J, Nathaniel J. 2020a. Evolution of human-environmental interactions in China from the Late Paleolithic to the Bronze Age. Prog Phys Geogr-Earth Environ, 44: 233–250
Dong G H, Liu F W, Yang Y S, Wang L, Chen F H. 2016. Cultural expansion and its influencing factors during Neolithic period in the Yellow River valley, northern China (in Chinese). Chin J Nat, 38: 248–252
Dong G H, Yang Y S, Han J Y, Wang H, Chen F H. 2017. Exploring the history of cultural exchange in prehistoric Eurasia from the perspectives of crop diffusion and consumption. Sci China Earth Sci, 60: 1110–1123
Dong G H, Yang Y S, Liu X Y, Li H M, Cui Y F, Wang H, Chen G K, Dodson J, Chen F H. 2018. Prehistoric trans-continental cultural exchange in the Hexi Corridor, northwest China. Holocene, 28: 621–628
Dong G H, Yang Y S, Ren L L, Ma M M. 2020c. The Human Subsistence Strategy Patterns and Human-Environment Interactions in Prehistoric Hexi Corridor (in Chinese). Beijing: Science Press. 78–96
Dong G H. 2018. A new story for wheat into China. Nat Plants, 4: 243–244
Doppler T, Pichler S, Röder B, Schibler J. 2013. Coping with crises I: Subsistence variety and resilience in the late Neolithic lakeshore settlement Arbon Bleiche 3 (Switzerland). In: Kerig T, Zimmermann A, eds. Economic Archaeology: Form Structure to Performance in European Archaeology. Bonn: Hablet. 163–174
Doumani P N, Frachetti M D, Beardmore R, Schmaus T M, Spengler R NIII, Mar’yashev A N. 2015. Burial ritual, agriculture, and craft production among Bronze Age pastoralists at Tasbas (Kazakhstan). Archaeol Res Asia, 1–2: 17–32
Du L Y, Ma M M, Lu Y W, Dong J J, Dong G H. 2020. How did human activity and climate change influence animal exploitation during 7500–2000 BP in the Yellow River Valley, China? Front Ecol Evol, 8: 161
Editorial Board of General Annals of Chinese Culture. 1998. Cultural Exchange between China and Central Asia (in Chinese). Shanghai: Shanghai People Press
Editorial Department of China Agricultural Encyclopedia. 1991. Encyclopedia of Chinese Agriculture (Crops Volume) (in Chinese). Beijing: Agriculture Press
Fages A, Hanghøj K, Khan N, Gaunitz C, Seguin-Orlando A, Leonardi M, McCrory Constantz C, Gamba C, Al-Rasheid K A S, Albizuri S, Alfarhan A H, Allentoft M, Alquraishi S, Anthony D, Baimukhanov N, Barrett J H, Bayarsaikhan J, Benecke N, Bernáldez-Sánchez E, Berrocal-Rangel L, Biglari F, Boessenkool S, Boldgiv B, Brem G, Brown D, Burger J, Crubézy E, Daugnora L, Davoudi H, de Barros Damgaard P, de los Ángeles de Chorro y de Villa-Ceballos M, Deschler-Erb S, Detry C, Dill N, do Mar Oom M, Dohr A, Ellingvåg S, Erdenebaatar D, Fathi H, Felkel S, Fernández-Rodríguez C, García-Viñas E, Germonpré M, Granado J D, Hallsson J H, Hemmer H, Hofreiter M, Kasparov A, Khasanov M, Khazaeli R, Kosintsev P, Kristiansen K, Kubatbek T, Kuderna L, Kuznetsov P, Laleh H, Leonard J A, Lhuillier J, Liesau von Lettow-Vorbeck C, Logvin A, Lõugas L, Ludwig A, Luis C, Arruda A M, Marques-Bonet T, Matoso Silva R, Merz V, Mijiddorj E, Miller B K, Monchalov O, Mohaseb F A, Morales A, Nieto-Espinet A, Nistelberger H, Onar V, Pálsdóttir A H, Pitulko V, Pitskhelauri K, Pruvost M, Rajic Sikanjic P, Rapan Papeša A, Roslyakova N, Sardari A, Sauer E, Schafberg R, Scheu A, Schibler J, Schlumbaum A, Serrand N, Serres-Armero A, Shapiro B, Sheikhi Seno S, Shevnina I, Shidrang S, Southon J, Star B, Sykes N, Taheri K, Taylor W, Teegen W R, Trbojević Vukičević T, Trixl S, Tumen D, Undrakhbold S, Usmanova E, Vahdati A, Valenzuela-Lamas S, Viegas C, Wallner B, Weinstock J, Zaibert V, Clavel B, Lepetz S, Mashkour M, Helgason A, Stefánsson K, Barrey E, Willerslev E, Outram A K, Librado P, Orlando L. 2019. Tracking five millennia of horse management with extensive ancient genome time series. Cell, 177: 1419–1435.e31
Fairbairn A, Asouti E, Near J, Martinoli D. 2002. Macro-botanical evidence for plant use at Neolithic Çatalhöyük south-central Anatolia, Turkey. Veget Hist Archaeobot, 11: 41–54
Fairbairn A, Near J, Martinoli D. 2005. Macrobotanical investigations of the North, South and KOPAL areas at Catalhoyuk. In: Hodder I, ed. Inhabiting Çatalhöyük: Reports from the 1995–1999 Seasons. Cambridge: McDonald Institute for Archaeological Research. 137–201
Frachetti M D, Smith C E, Traub C M, Williams T. 2017. Nomadic ecology shaped the highland geography of Asia’s silk roads. Nature, 543: 193–198
Fu Q M, Jin S A, Hu Y W, Ma Z, Pan J C, Wang C S. 2010. Agricultural development and human diets in Gouwan site, Xichuan, Henan (in Chinese). Chin Sci Bull, 55: 589–595
Gansu Provincial Institute of Cultural Relics and Archaeology. 2006. Dadiwan in Qin’an: Report on Excavations at a Neolithic Site (in Chinese). Beijing: Cultural Relics Publishing House. 693–694
Gaunitz C, Fages A, Hanghøj K, Albrechtsen A, Khan N, Schubert M, Seguin-Orlando A, Owens I J, Felkel S, Bignon-Lau O, de Barros Damgaard P, Mittnik A, Mohaseb A F, Davoudi H, Alquraishi S, Al-farhan A H, Al-Rasheid K A S, Crubézy E, Benecke N, Olsen S, Brown D, Anthony D, Massy K, Pitulko V, Kasparov A, Brem G, Hofreiter M, Mukhtarova G, Baimukhanov N, Lõugas L, Onar V, Stockhammer P W, Krause J, Boldgiv B, Undrakhbold S, Erdenebaatar D, Lepetz S, Mashkour M, Ludwig A, Wallner B, Merz V, Merz I, Zaibert V, Willerslev E, Librado P, Outram A K, Orlando L. 2018. Ancient genomes revisit the ancestry of domestic and Przewalski’s horses. Science, 360: 111–114
Gignoux C R, Henn B M, Mountain J L. 2011. Rapid, global demographic expansions after the origins of agriculture. Proc Natl Acad Sci USA, 108: 6044–6049
Gong W. 2016. Research of human diet in Shang Period of the Daxinzhuang site and Liujiazhuang site: Results from the plant remains and the stable isotope of carbon and nitrogen (in Chinese). Master’s Dissertation. Ji’nan: Shandong University. 1–127
Greenfield H J, Arnold E R. 2015. New perspectives on the zooarchaeological evidence for the earliest intensification of dairying in south eastern Europe. World Archaeol, 47: 792–818
Greenfield H J, Greenfield T L J. 2014. Subsistence and settlement in the early Neolithic of temperate SE Europe: A view from Blagotin, Serbia. Archaeol Bul, 1: 1–33
Guan Y, Gao X, Li F, Pei S W, Chen F Y, Zhou Z Y. 2012. Modern human behaviors during the late stage of the MIS3 and the broad spectrum revolution: Evidence from a Shuidonggou Late Paleolithic site. Chin Sci Bull, 57: 379–386
Guo Y, Hu Y W, Gao Q, Wang Q S, Richards M P. 2011. Stable carbon and nitrogen isotope evidence in human diets based on evidence from the Jiangzhai site, China (in Chinese). Acta Anthropol Sin, 30: 149–157
Guo Y, Xia Y, Dong Y F, Yu B Y, Fan Y L, Wen F Y, Gao Q. 2016. Stable isotope analysis of human at Beiliu site (in Chinese). Archaeol Cult Relics, 1: 115–120
Han J Y. 2021. The three stages of early Sino-western cultural interactions (in Chinese). Acta Archaeol Sin, 3: 317–338
Hanks B K, Linduff K M. 2009. Social Complexity in Prehistoric Eurasia: Monuments, Metals and Mobility. Cambridge: Cambridge University Press
Harmatha J. 2002. History of Civilizations of Central Asia II (Chinese Edition) (in Chinese). Beijing: China Translation and Publishing Corporation. 142–157
Harris D R, Masson V M, Berezkin Y E, Charles M P, Gosden C, Hillman G C, Kasparov A K, Korobkova G F, Kurbansakhatov K, Legge A J, Limbrey S. 1993. Investigating early agriculture in Central Asia: New research at Jeitun, Turkmenistan. Antiquity, 67: 324–338
Henan Provincial Institute of Cultural Relics and Archaeology. 2009. Sanmenxia Nanjiaokou (in Chinese). Beijing: Science Press. 316–320
Hermes T R, Tishkin A A, Kosintsev P A, Stepanova N F, Krause-Kyora B, Makarewicz C A. 2020. Mitochondrial DNA of domesticated sheep confirms pastoralist component of Afanasievo subsistence economy in the Altai Mountains (3300–2900 cal BC). Archaeol Res Asia, 24: 100232
Hovsepyan R, Willcox G. 2008. The earliest finds of cultivated plants in Armenia: Evidence from charred remains and crop processing residues in pisé from the Neolithic settlements of Aratashen and Aknashen. Veget Hist Archaeobot, 17: 63–71
Hovsepyan R. 2013. First archaeobotanical data from the basin of Lake Sevan. In: Archäologie in Armenien II, eds. Veröffentlichungen des Landesamtes für Denkmalpflege und Archäologie Sachsen-Anhalt 67. Halle (Saale): Landesmuseum für Vorgeschichte. 93–105
Hrisrova I, Atanassova J, Marinova E. 2016. Plant economy and vegetation of the Iron Age in Bulgaria: Archaeobotanical evidence from pit deposits. Archaeol Anthropol Sci, 9: 1481–1494
Hu S M, Yang M M, Sun Z Y, Shao J. 2016. Research on faunal remains from the 2012–2013 season excavation at the Shimao site in Shenmu, Shaanxi Province (in Chinese). Archaeol Cult Relics, 4: 109–121
Hu S M, Yang Q H, Yang M M. 2011. Analysis on the faunal remains from the Xinlefang site, Huayin, Shaanxi Province (in Chinese). Archaeol Cult Relics, 6: 117–125
Hu Y W, Luan F S, Wang S G, Wang C S, Richards M P. 2009. Preliminary attempt to distinguish the domesticated pigs from wild boars by the methods of carbon and nitrogen stable isotope analysis. Sci China Ser D-Earth Sci, 52: 85–92
Hu Y W, Ambrose S H, Wang C S. 2007. Stable isotopic analysis on ancient human bones in Jiahu site. Sci China Ser D-Earth Sci, 50: 563–570
Institute of Archaeology, CASS. 1999. Shizhaocun and Xishanping (in Chinese). Beijing: Encyclopedia of China Publishing House
Ji D X, Chen F H, Bettinger R L, Elston R G, Geng Z Q, Barton L, Wang H, An C B, Zhang D J. 2005. Human response to the Last Glacial Maximum: Evidence from North China (in Chinese). Acta Anthropol Sin, 24: 270–282
Jia X. 2012. Cultural evolution process and plant remains during Neolithic-Bronze Age in northeast Qinghai Province (in Chinese). Doctoral Dissertation. Lanzhou: Lanzhou University. 47–56
Jiang H, Wu Y, Wang H, Ferguson D K, Li C S. 2013. Ancient plant use at the site of Yuergou, Xinjiang, China: Implications from desiccated and charred plant remains. Veget Hist Archaeobot, 22: 129–140
Jin G Y, Wang H Y, Yan S D, Liu C J, Lan Y F, Tong P H. 2011. Study on the carbonized plant remains at the site during Longshan Culture of Zhaojiazhuang in Jiaozhou, Shandong Province (in Chinese). In: Center for Science and Technology Archaeology, Institute of Archaeology, CASS, eds. Science for Archaeology (III). Beijing: Science Press. 36–53
Jones M K, Hunt H, Lightfoot E, Lister D, Liu X Y, Matuzeviciute G M. 2011. Food globalization in prehistory. World Archaeol, 43: 665–675
Kamjan S, Mashkour M, Mohaseb F A, Fathi H, Azizi K H. 2018. Zooarchaeology of the Pre-Pottery and Pottery Neolithic site of Qasr-e Ahmad (Iran). In: Çakirlar C, Chahoud J, Berthon R, Birch S P, eds. Archaeozoology of the Near East XII. Groningen: Barkhuis Publishing. 27–40
Kuijt I, Goring-Morris N. 2002. Foraging, farming, and social complexity in the Pre-Pottery Neolithic of the southern Levant: A review and synthesis. J World Prehistory, 16: 361–440
Lecavalier B S, Fisher D A, Milne G A, Vinther B M, Tarasov L, Huybrechts P, Lacelle D, Main B, Zheng J, Bourgeois J, Dyke A S. 2017. High Arctic Holocene temperature record from the Agassiz ice cap and Greenland ice sheet evolution. Proc Natl Acad Sci USA, 114: 5952–5957
Lee G A, Crawford G W, Liu L, Chen X. 2007. Plants and people from the Early Neolithic to Shang periods in North China. Proc Natl Acad Sci USA, 104: 1087–1092
Lelli R, Allen R, Biondi G, Calattini M, Barbaro C C, Gorgoglione M A, Manfredini A, Martínez-Labarga C, Radina F, Silvestrini M, Tozzi C, Rickards O, Craig O E. 2012. Examining dietary variability of the earliest farmers of South-Eastern Italy. Am J Phys Anthropol, 149: 380–390
Li J Y, Ilvonen L, Xu Q H, Ni J, Jin L Y, Holmström L, Zheng Z, Lu H Y, Luo Y L, Li Y C, Li C H, Zhang X J, Seppä H. 2015. East Asian summer monsoon precipitation variations in China over the last 9500 years: A comparison of pollen-based reconstructions and model simulations. Holocene, 26: 592–602
Li X Q, Zhou X Y, Zhou J, Dodson J, Zhang H B, Shang X. 2007. The earliest archaeobiological evidence of the broadening agriculture in China recorded at Xishanping site in Gansu Province. Sci China Ser D-Earth Sci, 50: 1707–1714
Li Y C, Tian J Y, Liu F W, Yang B Y, Gu K S Y, Rahman Z U, Yang L Q, Chen F H, Dong G H, Kong Q P. 2019. Neolithic millet farmers contributed to the permanent settlement of the Tibetan Plateau by adopting barley agriculture. Natl Sci Rev, 6: 1005–1013
Li Y, Zhang C R, Taylor W T T, Chen L, Flad R K, Boivin N, Liu H, You Y, Wang J X, Ren M, Xi T Y, Han Y F, Wen R, Ma J. 2020. Early evidence for mounted horseback riding in northwest China. Proc Natl Acad Sci USA, 117: 29569–29576
Li Z P, Ren L L, Shi Y Z. 2015. Identification and preliminary study of the animal remains unearthed in Xiaoli site, Xingtai City, Hebei Province (in Chinese). Cult Relics South China, 4: 92–119
Lillie M, Budd C, Potekhina I. 2011. Stable isotope analysis of prehistoric populations from the cemeteries of the Middle and Lower Dnieper Basin, Ukraine. J Archaeol Sci, 38: 57–68
Lin M H, Luan F S, Fang H, Xu H, Zhao H T, Barker G. 2018. Pathological evidence reveals cattle traction in North China by the early second millennium BC. Holocene, 28: 1205–1215
Lin M H. 2011. Analyses on faunal remains from Longshangang Site, in Xichuan County, Henan Province (in Chinese). Master’s Dissertation. Ji’nan: Shandong University. 1–133
Lister D L, Jones M K. 2013. Is naked barley an eastern or a western crop? The combined evidence of archaeobotany and genetics. Veget Hist Archaeobot, 22: 439–446
Liu C J, Jin G Y, Kong Z C. 2008. Paleoethnobotany: Research on Seeds and Fruits (in Chinese). Beijing: Science Press. 168–170
Liu C J, Kong Z C, Lang S D. 2004. Relics of crops and discussion on the environment of primitive people living in Dadiwan site (in Chinese). Cul Rel Central China, 4: 26–30
Liu C Z. 2003. The vulnerability of water resources in Northwest China. J Glaciol Geocryol, 25: 309–314
Liu H. 2019. The research of animal remains of Maojiaping site in Tianshui, Gansu Province (in Chinese). Doctoral Dissertation. Xi’an: Northwest Univeristy. 1–191
Liu X Y, Jones P J, Matuzeviciute G M, Hunt H V, Lister D L, An T, Przelomska N, Kneale C J, Zhao Z, Jones M K. 2019. From ecological opportunism to multi-cropping: Mapping food globalisation in prehistory. Quat Sci Rev, 206: 21–28
Liu X Y, Lightfoot E, O’Connell T C, Wang H, Li S C, Zhou L P, Hu Y W, Motuzaite-Matuzeviciute G, Jones M K. 2014. From necessity to choice: Dietary revolutions in west China in the second millennium BC. World Archaeol, 46: 661–680
Liu X Y, Lister D L, Zhao Z J, Staff R A, Jones P J, Zhou L P, Pokharia A K, Petrie C A, Pathak A, Lu H L, Matuzeviciute G M, Bates J, Pilgram T K, Jones M K. 2016. The virtues of small grain size: Potential pathways to a distinguishing feature of Asian wheats. Quat Int, 426: 107–119
Liu X Y. 2014. Study on the plant remains excavated in Anban site in 2012 (in Chinese). Master’s Dissertation. Xi’an: Northwest University. 1–60
Long T W, Leipe C, Jin G Y, Wagner M, Guo R Z, Schroder O, Tarasov P E. 2018. The early history of wheat in China from 14C dating and Bayesian chronological modelling. Nat Plants, 4: 272–279
Lösch S, Grupe G, Peters J. 2006. Stable isotopes and dietary adaptations in humans and animals at pre-pottery Neolithic Nevalli Çori, southeast Anatolia. Am J Phys Anthropol, 131: 181–193
Lu H Q, Zhou C W. 1990. Identification report of animal and plant specimens unearthed at Yinjiacheng Site in Sichui County, Shandong Province. In: Department of History, Shandong University, eds. Sishui Yinjiacheng. Beijing: Cultural Relics Press. 350–352
Ma M M, Dong G H, Jia X, Wang H, Cui Y F, Chen F H. 2016. Dietary shift after 3600 cal yr BP and its influencing factors in northwestern China: Evidence from stable isotopes. Quat Sci Rev, 145: 57–70
Manning K, Downey S S, Colledge S, Conolly J, Stopp B, Dobney K, Shennan S. 2013. The origins and spread of stock-keeping: The role of cultural and environmental influences on early Neolithic animal exploitation in Europe. Antiquity, 87: 1046–1059
Manning K, Timpson A, Colledge S, Crema E, Shennan S. 2015. The Cultural Evolution of Neolithic Europe. EUROEVOL Dataset
Manning K. 2016. The cultural evolution of Neolithic Europe. EUROEVOL Dataset 2: Zooarchaeological data. J Open Archaeol Data, 5: E3
Marcott S A, Shakun J D, Clark P U, Mix A C. 2013. A reconstruction of regional and global temperature for the past 11,300 years. Science, 339: 1198–1201
Mileto S, Kaiser E, Rassamakin Y, Whelton H, Evershed R P. 2017. Differing modes of animal exploitation in North-Pontic Eneolithic and Bronze Age societies. STAR-Sci Tech Archaeol Res, 3: 112–125
Miller A R V, Makarewicz C A. 2019. Intensification in pastoralist cereal use coincides with the expansion of trans-regional networks in the Eurasian Steppe. Sci Rep, 9: 8363
Miller N F, Spengler R N, Frachetti M. 2016. Millet cultivation across Eurasia: Origins, spread, and the influence of seasonal climate. Holocene, 26: 1566–1575
Miyake H, Hiroshi M, Maeda O, Tada S, Homgo K, Dano K, Yoshida K. 2009. Archaeological site of Salat Jermy Janu (Republic of Turkey) Overview: 2004–2008 (in Japanese). Prehist Archaeol Res, 20: 75–112
Moore A M T, Hillman G C, Legge A J. 1975. The excavation of Tell Abu Hureyra in Syria: A preliminary report. Proc Prehist Soc, 41: 50–77
Motuzaite-Matuzeviciute G, Kiryushin Y F, Rakhimzhanova S Z, Svyatko S, Tishkin A A, O’Connell T C. 2016. Climatic or dietary change? Stable isotope analysis of Neolithic-Bronze Age populations from the Upper Ob and Tobol River basins. Holocene, 26: 1711–1721
Motuzaite-Matuzeviciute G, Staff R A, Hunt H V, Liu X Y, Jones M K. 2013. The early chronology of broomcorn millet (Panicum miliaceum) in Europe. Antiquity, 87: 1073–1085
Motuzaite-Matuzeviciute G, Telizhenko S, Jones M K. 2012. Archaeobotanical investigation of two Scythian-Sarmatian period pits in eastern Ukraine: Implications for floodplain cereal cultivation. J Field Archaeol, 37: 51–60
Nagasawa K. 1994. Silk Road and East-West cultural communications in the ancient Eurasia. In: Zhang Z Y, ed. Steppe Silk Road and Central Asian Civilization (in Chinese). Urumqi: Xinjiang Fine Arts Photography Publishing House. 310–314
Ning C, Wang C C, Gao S Z, Yang Y, Zhang X, Wu X Y, Zhang F, Nie Z Z, Tang Y P, Robbeets M, Ma J, Krause J, Cui Y Q. 2019. Ancient genome reveal Yamnaya-related ancestry and a potential source of Indo-European speakers in Iron Age Tianshan. Curr Biol, 29: 2526–2532.e4
Nomokonova T, Losey R J, Goriunova O I, Novikov A G, Weber A W. 2015. A 9,000 year history of seal hunting on Lake Baikal, Siberia: The zooarchaeology of Sagan-Zaba II. PLoS ONE, 10: e0128314
Oelze V M, Siebert A, Nicklisch N, Meller H, Dresely V, Alt K W. 2011. Early Neolithic diet and animal husbandry: Stable isotope evidence from three Linearbandkeramik (LBK) sites in Central Germany. J Archaeol Sci, 38: 270–279
Outram A K, Kasparov A, Stear N A, Varfolomeev V, Usmanova E, Evershed R P. 2012. Patterns of pastoralism in later Bronze Age Kazakhstan: New evidence from faunal and lipid residue analyses. J Archaeol Sci, 39: 2424–2435
Outram A K, Stear N A, Bendrey R, Olsen S, Kasparov A, Zaibert V, Thorpe N, Evershed R P. 2009. The earliest horse harnessing and milking. Science, 323: 1332–1335
Outram A K, Stear N A, Kasparov A, Usmanova E, Varfolomeev V, Evershed R P. 2011. Horses for the dead: Funerary foodways in Bronze Age Kazakhstan. Antiquity, 85: 116–128
Pechenkina E A, Ambrose S H, Ma X L, BenferJr R A. 2005. Reconstructing northern Chinese Neolithic subsistence practices by isotopic analysis. J Archaeol Sci, 32: 1176–1189
Perlès C, Quiles A, Valladas H. 2013. Early seventh-millennium AMS dates from domestic seeds in the initial Neolithic at Franchthi Cave (Argolid, Greece). Antiquity, 87: 1001–1015
Qi G Q, Lin Z Y, An J Y. 2006. Report of identification of faunal remains from the Dadiwan site. In: Gansu Provincial Institute of Cultural Relics and Archaeology, ed. Qin’an Dadiwan (in Chinese). Beijing: Cultural Relics Press. 861–910
Qinghai Provincial Archaeology Team. 1979. No.1 tomb of Majiayao culture in Hetaozhuang, Minhe county, Qinghai Province (in Chinese). Cult Relics, 9: 29–32
Qu Y T, Hu X J, Wang T T, Yang Y M. 2020. Early interaction of agropastoralism in Eurasia: New evidence from millet-based food consumption of Afanasyevo humans in the southern Altai Mountains, Xinjiang, China. Archaeol Anthropol Sci, 12: 195
Rassamakin Y. 1999. The neolithic of the Black Sea steppe: Dynamics of cultural and economic development 4500–2300 B.C. In: Levine M, Rassamakin Y, Kislenko A, Tatarintseva N, eds. Late Prehistoric Exploitation of the Eurasian Steppe. Cambridge: McDonald Institute Monographs. 59–182
Rawson J. 2017. China and the steppe: Reception and resistance. Antiquity, 91: 375–388
Redding R W. 2003. First report on faunal remains. In: Alizadeh A, eds. Excavations at the Prehistoric Mound of Chogha Bonut, Khuzestan, Iran (Seasons 1976/77, 1977/78, and 1996). Michigan: McNaughton & Gunn, Saline. 137–147
Reich D. 2018. Who We Are and How We Got Here: Ancient DNA and the New Science of the Human Past. Oxford: Oxford University Press
Ren L L. 2017. A study on animal exploitation strategies from the late Neolithic to Bronze Age in northeastern Tibetan Plateau and its surrounding areas, China (in Chinese). Doctoral Dissertation. Lanzhou: Lanzhou University. 1–151
Richards M P, Pearson J A, Molleson T I, Russell N, Martin L. 2003. Stable isotope evidence of diet at Neolithic Çatalhöyük, Turkey. J Archaeol Sci, 30: 67–76
Riehl S, Zeidi M, Conard N J. 2013. Emergence of agriculture in the foothills of the Zagros Mountains of Iran. Science, 341: 65–67
Roffet-Salque M, Marciniak A, Valdes P J, Pawlowska K, Pyzel J, Czerniak L, Krüger M, Roberts C N, Pitter S, Evershed R P. 2018. Evidence for the impact of the 8.2-ky BP climate event on Near Eastern early farmers. Proc Natl Acad Sci USA, 115: 8705–8709
Shaanxi Provincial Institute of Archaeology, Weinan Municipal Administration of Cultural Relics and Tourism, Hua County Administration of Cultural Relics and Tourism. 2014. The Quanhucun Site in Huaxian County-A Report of the 1997-Year Excavation (in Chinese). Beijing: Cultural Relics Press. 595–661
Shishlina N I, van der Plicht J, Hedges R E M, Zazovskaya E P, Sevastyanov V S, Chichagova O A. 2007. The catacomb cultures of the northwest Caspian steppe: 14C chronology, reservoir effect, and paleodiet. Radiocarbon, 49: 713–726
Song Y B, Li Q, He D L. 2013. The faunal remains analysis of Houyangguanzhuang site, Cangshan County, Shandong Province (in Chinese). In: Shandong Provincial Institute of Cultural Relics and Archaeology, ed. Haidai Archaeology (VI). Beijing: Science Press. 108–132
Song Y B. 2008. Analysis of animal remains unearthed in 2003 from Yuezhuang site, Changqing District, Jinan City (in Chinese). Archaeol Res, 7: 519–531
Spengler R NIII, Chang C, Tourtellotte P A. 2013. Agricultural production in the Central Asian mountains: Tuzusai, Kazakhstan (410–150 BC). J Field Archaeol, 38: 68–85
Spengler R NIII, de Nigris I, Cerasetti B, Carra M, Rouse L M. 2018. The breadth of dietary economy in Bronze Age Central Asia: Case study from Adji Kui 1 in the Murghab region of Turkmenistan. J Archaeol Sci-Rep, 22: 372–381
Spengler R NIII, Miller N F, Neef R, Tourtellotte P A, Chang C. 2017. Linking agriculture and exchange to social developments of the central Asian Iron Age. J Anthropol Archaeol, 48: 295–308
Spengler R NIII. 2015. Agriculture in the central Asian Bronze Age. J World Prehist, 28: 215–253
Spengler R, Frachetti M, Doumani P, Rouse L, Cerasetti B, Bullion E, Mar’yashev A. 2014. Early agriculture and crop transmission among Bronze Age mobile pastoralists of Central Eurasia. Proc R Soc B, 281: 20133382
Staubwasser M, Weiss H. 2006. Holocene climate and cultural evolution in late prehistoric-early historic West Asia. Quat Res, 66: 372–387
Stevens C J, Murphy C, Roberts R, Lucas L, Silva F, Fuller D Q. 2016. Between China and South Asia: A Middle Asian corridor of crop dispersal and agricultural innovation in the Bronze Age. Holocene, 26: 1541–1555
Suo X F, Li S B. 2013. Neolithic animal remains in Liaoxi region (in Chinese). Steppe Cult Relics, 1: 43–49, 115
Svyatko S V, Schulting R J, Mallory J, Murphy E M, Reimer P J, Khartanovich V I, Chistov Y K, Sablin M V. 2013. Stable isotope dietary analysis of prehistoric populations from the Minusinsk Basin, southern Siberia, Russia: A new chronological framework for the introduction of millet to the Eastern Eurasian Steppe. J Archaeol Sci, 40: 3936–3945
Tan L C, Dong G H, An Z S, Edwards R L, Li H M, Li D, Spengler R, Cai Y J, Cheng H, Lan J H, Orozbaev R, Liu R L, Chen J H, Xu H, Chen F H. 2021. Megadrought and cultural exchange along the proto-silk road. Sci Bull, 66: 603–611
Tian D, Festa M, Cong D X, Zhao Z J, Jia P W, Betts A. 2021b. New evidence for supplementary crop production, foddering and fuel use by Bronze Age transhumant pastoralists in the Tianshan Mountains. Sci Rep, 11: 13718
Tian D, Ma J, Ren M, Xi T Y, Wang J X, Zhao Z J. 2021a. Barley production in early Xinjiang: Evidence from archaeobotanical remains in the Shirenzigou site, Northern Tianshan Mountains (in Chinese). Agricult His China, 3: 44–55
Tong E Z. 1986. On the borderland half-moon cultural transmission zone from northeast to southwest of China (in Chinese). Proc Cult Relics Archaeol. 17–43
Vaiglova P, Reid R E B, Lightfoot E, Pilaar Birch S E, Wang H, Chen G, Li S, Jones M, Liu X. 2021. Localized management of non-indigenous animal domesticates in northwestern China during the Bronze Age. Sci Rep, 11: 15764
Verdugo M P, Mullin V E, Scheu A, Mattiangeli V, Daly K G, Maisano Delser P, Hare A J, Burger J, Collins M J, Kehati R, Hesse P, Fulton D, Sauer E W, Mohaseb F A, Davoudi H, Khazaeli R, Lhuillier J, Rapin C, Ebrahimi S, Khasanov M, Vahidi S M F, MacHugh D E, Ertugrul O, Koukouli-Chrysanthaki C, Sampson A, Kazantzis G, Kontopoulos I, Bulatovic J, Stojanovic I, Mikdad A, Benecke N, Linstädter J, Sablin M, Bendrey R, Gourichon L, Arbuckle B S, Mashkour M, Orton D, Horwitz L K, Teasdale M D, Bradley D G. 2019. Ancient cattle genomics, origins, and rapid turnover in the Fertile Crescent. Science, 365: 173–176
Wang C C, Reinhold S, Kalmykov A, Wissgott A, Brandt G, Jeong C, Cheronet O, Ferry M, Harney E, Keating D, Mallick S, Rohland N, Stewardson K, Kantorovich A R, Maslov V E, Petrenko V G, Erlikh V R, Atabiev B C, Magomedov R G, Kohl P L, Alt K W, Pichler S L, Gerling C, Meller H, Vardanyan B, Yeganyan L, Rezepkin A D, Mariaschk D, Berezina N, Gresky J, Fuchs K, Knipper C, Schiffels S, Balanovska E, Balanovsky O, Mathieson I, Higham T, Berezin Y B, Buzhilova A, Trifonov V, Pinhasi R, Belinskij A B, Reich D, Hansen S, Krause J, Haak W. 2019. Ancient human genome-wide data from a 3000-year interval in the Caucasus corresponds with eco-geographic regions. Nat Commun, 10: 590
Wang C C, Yeh H Y, Popov A N, Zhang H Q, Matsumura H, Sirak K, Cheronet O, Kovalev A, Rohland N, Kim A M, Mallick S, Bernardos R, Tumen D, Zhao J, Liu Y C, Liu J Y, Mah M, Wang K, Zhang Z, Adamski N, Broomandkhoshbacht N, Callan K, Candilio F, Carlson K S D, Culleton B J, Eccles L, Freilich S, Keating D, Lawson A M, Mandl K, Michel M, Oppenheimer J, Özdogan K T, Stewardson K, Wen S, Yan S, Zalzala F, Chuang R, Huang C J, Looh H, Shiung C C, Nikitin Y G, Tabarev A V, Tishkin A A, Lin S, Sun Z Y, Wu X M, Yang T L, Hu X, Chen L, Du H, Bayarsaikhan J, Mijiddorj E, Erdenebaatar D, Iderkhangai T O, Myagmar E, Kanzawa-Kiriyama H, Nishino M, Shinoda K I, Shubina O A, Guo J, Cai W, Deng Q, Kang L, Li D, Li D, Lin R, Nini, Shrestha R, Wang L X, Wei L, Xie G, Yao H, Zhang M, He G, Yang X, Hu R, Robbeets M, Schiffels S, Kennett D J, Jin L, Li H, Krause J, Pinhasi R, Reich D. 2021. Genomic insights into the formation of human populations in East Asia. Nature, 591: 413–419
Wang H Y, Liu Y C, Jin G Y. 2011. Analysis on the carbonized plant remains from the Dongpan Site, 2009, Linshu County, Shandong (in Chinese). In: Institute of Cultural Heritage, Shandong University, ed. East Asia Archaeology (VIII). Beijing: Science Press. 357–372, 455–459
Wang H, Wang W L, Hu S M, Louise M. 2013. The domestication and husbandry strategies of the pigs in the Wei River Valley during the Neolithic Age (in Chinese). Archaeology, 9: 87–98
Wang H. 2011. Animal subsistence of the Yangshao period in the Wei River Valley: A case-study from the site of Wayaogou in Shaanxi Province, China. Doctoral Dissertation. London: University College London
Wang T T, Wei D, Chang X, Yu Z Y, Zhang X Y, Wang C, Hu Y, Fuller B T. 2019. Tianshanbeilu and the isotopic millet road: Reviewing the late Neolithic/Bronze Age radiation of human millet consumption from north China to Europe. Natl Sci Rev, 6: 1024–1039
Wang X, Shang X, Jiang H E, Zhang P C, Wang W L, Wang C S. 2015. Preliminary analysis of flotation results of two sites in Baishuihe River Basin, Shaanxi Province (in Chinese). Archaeol Cult Relic, 2: 100–104
Watson A M. 1983. Agricultural Innovation in the Early Islamic World: The Diffusion of Crops and Farming Techniques, 700–1100. Cambridge: Cambridge University Press. 272
Weber A W, Link D W, Katzenberg M A. 2002. Hunter-gatherer culture change and continuity in the Middle Holocene of the Cis-Baikal, Siberia. J Anthropol Archaeol, 21: 230–299
Weber A W, White D, Bazaliiskii V I, Goriunova O I, Savel’ev N A, Anne Katzenberg M. 2011. Hunter-gatherer foraging ranges, migrations, and travel in the middle Holocene Baikal region of Siberia: Insights from carbon and nitrogen stable isotope signatures. J Anthropol Archaeol, 30: 523–548
Weninger B, Alram-Stern E, Bauer E, Clare L, Danzeglocke U, Jöris O, Kubatzki C, Rollefson G, Todorova H, van Andel T. 2006. Climate forcing due to the 8200 cal yr BP event observed at early Neolithic sites in the eastern Mediterranean. Quat Res, 66: 401–420
Willcox G, Stordeur D. 2012. Large-scale cereal processing before domestication during the tenth millennium cal BC in northern Syria. Antiquity, 86: 99–114
Willcox G. 2002. Geographical variation in major cereal components and evidence for independent domestication events in the Western Asia. In: Cappers R T J, Bottema S, eds. The Dawn of Farming in the Near East. Berlin: Ex Oriente. 133–140
Wu W W, Zhang K S, Wang Z B, Jin G Y. 2013. Analysis on the carbonized plant remains from the Xihe Site, 2008, Zhangqiu County, Shandong (in Chinese). In: Institute of Cultural Heritage, Shandong University, eds. East Asia Archaeology (X). Beijing: Science Press. 373–390, 477–478
Wu X, Miller N F, Crabtree P. 2015. Agro-pastoral strategies and food production on the achaemenid frontier in Central Asia: A case study of Kyzyltepa in southern Uzbekistan. Iran, 53: 93–117
Wu Z, Yuan J, Zhao X, Chen X L. 2016. The animal archaeology research on the domestic dogs unearthed from sites between Neolithic and Pre-Qin Period in China (in Chinese). Cult Relics Southern China, 3: 155–161
Yan W M. 2000. The Origin of Agriculture and Civilization (in Chinese). Beijing: Science Press. 50–89
Yang B, Qin C, Bräuning A, Osborn T J, Trouet V, Ljungqvist F C, Esper J, Schneider L, Grießinger J, Büntgen U, Rossi S, Dong G H, Yan M, Ning L, Wang J L, Wang X F, Wang S M, Luterbacher J, Cook E R, Stenseth N C. 2021. Long-term decrease in Asian monsoon rainfall and abrupt climate change events over the past 6,700 years. Proc Natl Acad Sci USA, 118: e2102007118
Yang C. 2007. The research of the animal remains of Xicha site in Neimenggu (in Chinese). Master’s Dissertation. Changchun: Jilin University. 1–77
Yang R P, Yang Y M, Li W Y, Abuduresule Y, Hu X J, Wang C S, Jiang H E. 2014. Investigation of cereal remains at the Xiaohe Cemetery in Xinjiang, China. J Archaeol Sci, 49: 42–47
Yang W Z, Yu C Z. 1992. Regional Governance and Evaluation of Loess Plateau (in Chinese). Beijing: Science Press. 73–124
Yang Y M, Shevchenko A, Knaust A, Abuduresule I, Li W Y, Hu X J, Wang C S, Shevchenko A. 2014. Proteomics evidence for kefir dairy in Early Bronze Age China. J Archaeol Sci, 45: 178–186
Yang Y S. 2017. The transition of human subsistence strategy and its influencing factors during prehistoric times in the Hexi Corridor, northwest China (in Chinese). Doctoral Dissertation. Lanzhou: Lanzhou University. 65–71
Yang Y Z, Hameed M, Sameer M A. 2020. Archaeobotanical insights of the pre-historic agriculture in Pakistan: Archaeological acumens. Ancient Punjab, 8: 86–102
Yatoo M A, Spate M, Betts A, Pokharia A K, Shah M A. 2020. New evidence from the Kashmir Valley indicates the adoption of East and West Asian crops in the western Himalayas by 4400 years ago. Quat Sci Adv, 2: 100011
Yu C, Lv P, Zhao C C. 2011. The identification and research of animal remains of Xishan site in Li County, Gansu Province (in Chinese). Cult Relics Southern China, 3: 73–79
Yu D. 2008. A study of the faunal remains from the Nanfangshui Site in Tang County, Hebei Province (in Chinese). Master’s Dissertation. Changchun: Jilin University. 1–54
Yuan J. 2010. Zooarchaeological study on the domestic animals in ancient China (in Chinese). Quat Sci, 30: 298–306
Zeder M A. 2008. Domestication and early agriculture in the Mediterranean Basin: Origins, diffusion, and impact. Proc Natl Acad Sci USA, 105: 11597–11604
Zeder M A, Hesse B. 2000. The initial domestication of goats (Capra hircus) in the Zagros Mountains 10000 years ago. Science, 287: 2254–2257
Zeist W V, Bakker-Heeres J A H. 1985. Archaeological studies in the Levant I. Neolithic sites in the Damascus basin: Aswad, Ghoraifé, Ramad. Palaeohistoria, 24: 165–256
Zeist W V, Roller G J. 1995. Plant remains from Asikli Höyük, a pre-pottery Neolithic site in central Anatolia. Veget Hist Archaebot, 4: 179–185
Zhang F, Ning C, Scott A, Fu Q M, Bjørn R, Li W Y, Wei D, Wang W J, Fan L Y, Abuduresule I, Hu X J, Ruan Q R, Niyazi A, Dong G H, Cao P, Liu F, Dai Q Y, Feng X T, Yang R W, Tang Z H, Ma P C, Li C X, Gao S Z, Xu Y, Wu S H, Wen S Q, Zhu H, Zhou H, Robbeets M, Kumar V, Krause J, Warinner C, Jeong C W, Cui Y Q. 2021. The genomic origins of the Bronze Age Tarim Basin mummies. Nature, 599: 256–261
Zhang G L, Wang S Z, Ferguson D K, Yang Y M, Liu X Y, Jiang H E. 2017. Ancient plant use and palaeoenvironmental analysis at the Gumugou Cemetery, Xinjiang, China: Implication from desiccated plant remains. Archaeol Anthropol Sci, 9: 145–152
Zhang Q C, Zhu H. 2011. Carbon and nitrogen stable isotope analysis of the human bones from the Gumugou cemetery in Xinjiang: A Preliminary exploration of the early population dietary in Lop Nur (in Chinese). Western Regions Studies, 3: 91–96
Zhang W X, Wang H. 2000. Study on ancient cultivated rice at Qingyang site in Gansu Province (in Chinese). Agricult Archaeol, 3: 80–85
Zhang X L, Qiu S H, Zhong J, Liang Z H. 2012. The carbon and nitrogen stable isotope analysis of human bone from Qianzhangda Cemetery in Tengzhou, Shangdong Province (in Chinese). Archaeology, 9: 83–96
Zhang X L, Qiu S H, Zhong J, Zhao X P, Sun F X, Cheng L Q, Guo Y Q, Lin X W, Ma X L. 2010. Studies on diet of the ancient people of the Yangshao cultural sites in the Central Plains (in Chinese). Acta Anthropol Sin, 29: 197–207
Zhang X L, Wang J X, Xian Z Q, Qiu S H. 2003. Research on the diet structure of ancient humans (in Chinese). Archaeology, 2: 158–171
Zhang X L, Zhang J, Li Z P, Zhang L R, Chen G K, Wang P, Wang H. 2015. A preliminary analysis to the diet of the primitive residents of the Xichengyi site in Zhangye, Gansu (in Chinese). Archaeology, 7: 110–120
Zhao C. 2009. A study on the faunal remains of prehistoric period in Guanzhong Region (in Chinese). Master’s Dissertation. Changchun: Jilin University. 1–48
Zhao K L, Li X Q, Zhou X Y, Dodson J, Ji M. 2013. Impact of agriculture on an oasis landscape during the late Holocene: Palynological evidence from the Xintala site in Xinjiang, NW China. Quat Int, 311: 81–86
Zhao Z J, Fang Y M. 2007. Flotation results and analysis of the Wang-chenggang site (in Chinese). Huaxia Archaeol, 2: 78–89
Zhao Z J, Zhang J Z. 2009. Flatation results of 2001 excation season from the Jiahu site (in Chinese). Archaeology, 8: 84–93
Zhao Z J, Zhao C H, Yu J C, Wang T, Cui T X, Guo J N. 2020. Flotation results and analysis of plant remains from Donghulin site in Beijing (in Chinese). Archaeology, 7: 99–106
Zhao Z J. 2014. The process of origin of agriculture in China: Archaeological evidence from flotation results (in Chinese). Quat Res, 34: 73–84
Zhou L G. 2017. Tombs and society of Longshan culture in Henan from the perspective of stable carbon and nitrogen isotopes (in Chinese). Huaxia Archaeol, 3: 145–152
Zhou X Y, Li X Q, Dodson J, Zhao K L. 2016. Rapid agricultural transformation in the prehistoric Hexi corridor, China. Quat Int, 426: 33–41
Zhou X Y, Yu J J, Spengler R N, Shen H, Li X Q. 2020. 5,200-year-old cereal grains from the eastern Altai Mountains redate the trans-Eurasian crop exchange. Nat Plants, 6: 78–87
Zhouyuan Archaeological Working Team. 2011. The excavation of the bronze casting remains in locus west of Zhuangli village at Zhouyuan site in the springs of 2003 and 2004 (in Chinese). Acta Archaeol Sin, 2: 245–316
Zuo H R. 2017. Review and prospect of the animal archaeology of the domestic sheep/goat in China (in Chinese). Cult Relics South China, 1: 155–163
Zuo X X, Lu H Y, Jiang L P, Zhang J P, Yang X Y, Huan X J, He K Y, Wang C, Wu N Q. 2017. Dating rice remains through phytolith carbon-14 study reveals domestication at the beginning of the Holocene. Proc Natl Acad Sci USA, 114: 6486–6491
Acknowledgements
This work was supported by the National Key R&D Program of China (Grant No. 2018YFA0606402), the Strategic Priority Research Program of the Chinese Academy of Sciences (A) (Grant No. XDA2004010101), and the Second Tibetan Plateau Scientific Expedition and Research Program (Grant No. 2019QZKK0601).
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Dong, G., Du, L., Yang, L. et al. Dispersal of crop-livestock and geographical-temporal variation of subsistence along the Steppe and Silk Roads across Eurasia in prehistory. Sci. China Earth Sci. 65, 1187–1210 (2022). https://doi.org/10.1007/s11430-021-9929-x
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DOI: https://doi.org/10.1007/s11430-021-9929-x